Prince Harry and Meghan Markle were largely cheered as they held hands and joined the rest of the Royal Family for a service at St Paul's Cathedral on the second day of the Queen's Platinum Jubilee celebrations.
It is the first time the couple are back in full public view alongside the rest of the Firm since they quit the monarchy for a new life in the US two years ago.
There was a loud reaction as the smiling couple arrived at the cathedral with some boos - but by the time they were greeted at the west door by the Dean of St Paul's cheers could be heard.
They spent a few minutes talking to the senior cleric before they entered the place of worship and proceeded to greet the long line of clergy who are taking part in today’s service of Thanksgiving.
Harry stopped to chat for a few moments with the Archbishop of York who later gave the sermon praising the Queen’s 70-year reign, while Meghan looked stunning in a Dior coat dress.
The couple's seats were in the second row of the congregation, with Princesses Beatrice and Eugenie and their husbands, and Lady Sarah Chatto, the daughter of Princess Margaret, and her family.
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They were seated behind the Earl and Countess of Wessex who were in the front row with their children, Lady Louise Mountbatten-Windsor and Viscount Severn, and the Duke and Duchess of Gloucester.
Across the aisle were Prince Charles and Camilla, who have ornate chairs. Alongside were the seats of the Duke and Duchess of Cambridge and the Princess Royal and her husband, Vice Admiral Tim Laurence.
They processed through the nave of the church hand in hand, with Meghan smiling and Harry biting his lip at times.
Harry and Meghan had to squeeze past his cousins Princess Beatrice and her husband Edoardo Mapelli Mozzi, and Princess Eugenie and her husband Jack Brooksbank, who were already seated, to get to their places.
Harry was spotted, with his mouth open, appearing to be enjoying a joke with another royal seated across the aisle.
However, they were not seen with the Queen as it was announced last night that she would not be attending the service due to "episodic mobility issues" and would instead watch the event at Windsor Castle. She was represented at the event by heir Prince Charles.
Yesterday, the pair were reunited with the royals at Horse Guards Parade for Trooping the Colour and were seen chatting and joking with some of the younger royals - although were not pictured with Kate or the other Cambridges.
Although there was no sign of their own children Archie and Lilibet, it has emerged that their daughter, who turns one tomorrow, had the chance to meet the Queen for the first time last night.
Omid Scobie, one of the authors of the Duke and Duchess of Sussex's biography Finding Freedom, told BBC Breakfast: "I think people are expecting some sort of big birthday extravagant event, that we're going to see photographs from. From what I'm told, we shouldn't expect anything.
"Those moments with Lilibet are very much private between them and the Queen and of course we know how much she's been looking forward to it.
"They've been held back by a pandemic. Of course the times that Harry has been here it's just been by himself for quite sombre occasions. And so this really was the first time.
"Of course we know the Queen went back to Windsor Castle yesterday, the couple went back to Windsor as well where they're staying at Frogmore Cottage.
"So that would have been the first moment or the first chance for her to meet her namesake."
He also said that Harry and Meghan hope to be as "low-profile as possible" during their trip to the UK for The Platinum Jubilee.
Mr Scobie added: "I spoke with people close to the couple... who said that they want to be as low-profile as possible during this trip.
"It's almost hard to believe, but I think yesterday Trooping the Colour was a great example. We didn't really catch sight of them on TV cameras."
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